- Received September 19, 2016.
- Accepted March 6, 2017.
- Published online April 10, 2017.
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© 2017. Published by The Company of Biologists Ltd http://creativecommons.org/licenses/by/3.0This is an Open Access article distributed under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/3.0), which permits unrestricted use, distribution and reproduction in any medium provided that the original work is properly attributed.
- 1School of Biological Sciences, Monash University, Clayton VIC 3800, Australia
- 2Current Address: School of Life Sciences, La Trobe University, Bundoora, VIC 3086, Australia
- 3Graduate School of Pharmaceutical sciences, Chiba University, Chuo-ku, 260-8675, Japan
- 4Department of Biological Science, Hiroshima University, Higashi-Hiroshima 722-0073, Japan
- 5Current Address: Department of Biochemistry and Molecular Biology, University of Melbourne, Parkville, VIC 3010, Australia
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